European Water Mite Research

A report on the water mite collection of the Zoological Institute of the Academy of Sciences, St. Petersburg
Reinhard Gerecke and Denis Tumanov
Introduction
During a meeting at St. Petersburg in the second half of August, 1997 (financed by a grant of the DFG) I had the occasion to produce a preliminary inventory of the water mites preserved in the collections of the Zoological Institute. The nucleus of this material is constituted by the collection Sokolow.
The composition of this part of the collection can be easily deduced from Sokolow's publications. In general, he preserved types of new species and other particularly interesting specimens on slides, the rest of the material studied in well-labeled glass tubes.
This is not the place to document presence/absence and status of types and other interesting specimens from his collection. However, as far as possible, reference is made to the presence of the designation "Type" on labels; contrariously, for the boxes 25/26 which contain nearly exclusively Type material, an asterisk behind the preparation number signs slides lacking the indication "Type". Designation of lectotypes and the interpretation of the real status of the slides now labelled as "Type" is planned for a future project in cooperation with Dr. D. Tumanov (St. Petersburg).
The following text aims to give a general idea on the dimensions and organization of this very important collection (including a tentative evaluation following the criteria of Mc Ginley, 1989). As detailed as possible, all additional, "non-Sokolow" material will be enumerated (gifts from other colleagues, partly determined or undetermined specimens from recent samplings). This documentation reveals numerous surprising details new to scientists working in aquatic acarology.

General state of the collection
Until the end of the 2nd world war, Sokolows collection was treated as his private possess, and was sorted following his own numeration system. Later, Sokolow gave his collection to the museum and the slides were marked with additional numbers referring to the museum inventory system. Numbers in red colour refer to the type catalogue conserved at the museum. These contradictory numbers may be one of the reasons for the partly disordinated present state of the slide collection that renders conservatory work difficulty. A further problem is that preparations which were produced during the past 40 decades by Sokolow, Jankowskaja (and also other specialists?) have been added to the collection in a somewhat unsystematic manner. For these reasons, the first work to be done for allowing easy access to the slide collection, was a new numeration of all boxes, together with the production of a complete list of their contents (part A of this report). If there are older labels on the boxes, they are cited in parentheses in order to ensure identification of the boxes. The new numbers given in 1997 differ from all other labelling in consisting of a point preceeding the number (.1, .2, ...). They are written with white ink.
As a difference to the slide collection, the fluid collection is perfectly organized in a first, geographic, and a second, systematic part. For this collection we give a list of its contents because it retains many interesting species (part B).

Evaluation of the collection after Mc Ginley (1989)

Level 1, conservation problems.

Only a few (<1 %) of the slides display "red flag" problems. A few cases of drying slide media or detaching labels were found. These problems could be resolved when the collection is brought to a definitive taxonomic systemation (see below). Some of the slides from recent collections display only provisory labelling, with collecting sites given as numbers only. A complete labelling of these preps should be done before they are integrated in the general collection. We should keep in mind that in such cases the actual position in certain boxes could be significant with regard to the geographical derivation of the animals.
Level 2, unidentified material, unsorted and effectively inaccessible to research community.

A large part (> 50%) of the liquid collection of Sokolow is not labelled on species level, often completely devoid of taxonomic labels. In most cases, it should be possible to understand the identity of the specimens from his collecting site lists (published and/or conserved at the museum). A prerequistite is being familar with Russian language and calligraphy. Thus, access to this part of the liquid collection is possible, but not easy and time-consuming.
Level 3, unidentified material, sorted and effectively accessible to research community.

In many cases, the material from recent, "post-Sokolow", collections is not even determined on family level. Where identifications on genus level are given, in several times they were found erroneous. However, this is a minor problem because the material in question is well sorted geographically and collections from different geographical areas should best be studied as a whole. "Hydracarina" can be considered a sufficient taxonomic category allowing access to the research community.
Level 4, identified material (to species level) not incorporated into general collection, and Level 5, inadequately cureated material, not meeting departmental standards.

No such material present at St. Petersburg
Level 6, Physical curation complete, meeting departmental standards

By far most of the slides and (with the restrictions given for Levels 3 and 4) the whole liquid collection fall within this category. But we should keep in mind that from a preliminary control of selected vial material results that genus level determinations are often wrong. Only after a thorough revision of the whole collection, the ratio of level 3, 4 and 6 material can seriously estimated.
Level 7, Physical curation complete, species level inventory complete.

This is true for about 50 % of the collection. There are complete lists for the collection Sokolow including a Type inventory, but these lists can be used only in part because: 1) many slides are out of place; 2) the correctness of all Type designations must be checked. As far as I could see, there is no inventory of the "post-Sokolow" collections (the following list gives a preliminar one).
Level 8 and Level 9, label data and research data captured in database

The relation consumed time/gained scientific information of capturing label and research data in a database would be low. Reaching these levels is not necessary.

Need for conservation measurements
As can be deduced from the following catalogue, the main problem is the taxonomic disorganization of the slide collection. Slides of one single species can be distributed in 10 different boxes.
I propose a complete integration of all slides in one single collection, sorted on family and genus level following the Viets catalogue (1987), or (on my opinion better:) sorted on genus level in alphabetical order. In any case, Sokolow's preps will be easily recogniseable from their characteristic labels. Therefore it is not necessary to separate them from the remaining material.
Further, I propose to integrate also all types into the general collection. A revision of the taxonomic status of all preparations labeled as "Type" is desireable. In many cases, the designation of lectotypes will be necessary.
During the reorganisation of the collection, all preps should be checked with regard to the quality of the imbedding fluid and eventual detaching labels.
A. SLIDE COLLECTION (BOX 1-34)
1. "Hydrachna"
1 Hydrachna geographica,
H. sp.,
H. unisetata paludosa
2 Hygrobates calliger 11
H. fluviatilis 4
indet. 1
3 Hydryphantidae indet.
Limnesia maculata 5
L. undulata 1
L. fulgida 1
Arrenurus sp. 2
Neumania spinipes 1
Lebertia lineata 1
Euthyas truncata 1
Hygrobates calliger 1
H. trigonicus 1
Atractides spinipes 2
Piona sp. Ny 1
4 Hygrobates fluviatilis 3
H. calliger 4
H. nigromaculatus 2
H. sp. 4
5 Atractides amplexus 4
A. anomalus 2
Atractides ap. 4
6 Piona uncata 1
P. longipalpis 2
P. coccinea 2
P. rotunda 2
P. stjördalensis 1
Limnesia maculata 1
L. polonica 1
Lebertia saxonica 1
L. sp. 1
Arrenurus sp. 1
Arrenurus tubulator 1
Hygrobates cf. intricatus 1

2. "Piona"
1 Piona sp. Dn 18
P. coccinea 1
2 Piona coccinoides 14
3 Piona coccinoides 1
P. variabilis dispersa 1
P. coccinea 2
P. conglobata 1
P. sp. Dn 1
P. aff. nodatoides 2
P. carnea 5
4 Piona paucipora 20
5 Piona paucipora 16
6 Piona rotundoides 7
7 Piona paucipora 13
P. variabilis 1
P. sp. Dn 1
8 Piona sp. Dn 7
P. nodata 1
Forelia variegator 1
Hygrobates foreli? 1

3. "Eylais"
1 Eylais aff. krendowskii 1
E. aff. soari 1
2 E. aff. spinipons ? 1
E. aff. meridionalis 1
E. sp. 2
E. aff. crassipalpis 1
E. aff. setosa 1
3 E. aff. discreta 2
4 E. hamata 19
5 E. (aff.) soari 9
6 E. soari 8
E. aff. aculeata 1
7 E. (aff.) rimsky-korssakowi 4
8 E. koenikei 2
9 E. setosa 4
10 E. extendens 17
11 E. degenerata 4
12 E. (aff.) rimosa 15
13 E. triarcuata 4
14 E. ussuriensis 6
15 E. undulosa 3
16 E. tantilla 3
E. tantilliformis 2
17 E. mülleri 7
E. mutila 2
18 E. tullgreni 1
19 E. (aff.) mülleri 17
20 E. (aff.) mülleri 18
21 E. infundibulifera 3
4. "Piona"
1 P. conglobata 18
2 P. carnea 8
3 P. coccinea 15
4 P. rotunda 16
5 P. clavicornis 1
6 P. nodatoides 5
7 P. nodata 15
dito, 1 larva
8 P. variabilis 1
9 P. coccinea 15
10 P. uncata 7
11 P. trisetica 1
12 P. coccinea 1
P. stjördalensis 8
13 P. stjördalensis 15
14 P. stjördalensis 1
P. disparilis 1
15 P. coccinea occulta
16 P. longipalpis 6
17 P. variabilis 6
18 P. uncata 9
5. "Lebertia"
1 Lebertia indet.
(Dn, Lv, Ad. Pilo-) 15
2 dito 18
3 L. exuta 1
4 L. exuta behningi 1
5 L. exuta ocaensis 9
6 L. dubia 1
7 L. stackelbergi stackelbergi 1
8 L. porosa 12
9 L. caucasica 3
L. insignis 1
10 L. lineata 5
11 L. insignis 1
12 L. zernovi 4
L. aff. saxonica 1
13 L. densa 4
14 L. ziabuschinskii 3
15 L. (aff.) ignatowi 1
L. caucasica 1
L. saxonica 4

6. "Midea, Mideopsis, Acalyptonotus, Arrenurus, Neumania"
1 Neumania deltoides 2
N. vernalis 2
Dn indet. 1
2 N. limosa 1
3 N. spinipes 4
4 N. vernalis laminifera 1
N. aff. vernalis 1
5 Midea orbiculata 2
6 Arrenurus sp. 2
Megaluracarus sp. 1
Arrenurus bicuspidator 1
A. aff. cuspidator 2
A. latus 1
A. maculator 3
Sperchon thori 1
7 Arrenurus zachariae 2
A. latus 1
A. membranata 1
A. aff. truncatellus 1
A. neumani 3
A. robustus 1
A. crassicaudatus ? 1
A. absurdus 1 (1/1/0)
A. aff. tricuspidator 1
8 Neumania deltoides 3
9 N. (aff.) deltoides 2
10 N. (aff.) deltoides 1
N. callosa 2
N. sp. indet. 5
11 Arrenurus latus 1
12 A. sinuator 2
13 Mideopsis orbicularis 12
14-18 empty
7. "Hygrobates"
1 Hygrobates fluviatilis 18
2 dito 16
3 H. octoporus 5 (3/2/0)
4 H. fluviatilis 8
H. fluviatilis berlesei 2
5 H. nigromaculatus 8
6 dito 18
7 H. trigonicus 15
8 dito 5
H. sp. indet. 13
9 dito 17
H. fluviatilis 1
10 H. nigromaculatus 1
H. aff. foreli 1
H. sp. 9
11 dito 18
12 H. calliger 18
13 H. (aff.) fluviatilis
(var. berlesei) 17
14 H. octoporus 12
15 H. longiporus 14
16 dito 10
17 H. longipalpis 17
18 H. calliger 15
19 H. kirgizicus 3
H. trigonicus 1
H. sp. 4
20 H. foreli 4 (2/2/0)
21 H. (aff.) foreli 16
8. varia, single specimens:
1 terrestrial mites (among them Discopama orchestiidarum, det. Oudemans)
2 Hydrochoreutes krameri
Brachypoda versicolor
Soldanellonyx monardi
Porolohmanella violacea
Piona coccinea
Limnesia undulata
3 Eylais tullgreni
E. infundibulifera
Neobrachypoda ekmani
Mideopsis crassipes
Arrenurus latus
A. aff. tubulator
A. maculator
A. zachariasi
Parathyas thoracata
Piona Lv on Chironomus
Sperchon Lv on Simuliidae
4 Piona, Unionicola, Eylais indet.
9. "Piona, Hydryphantes, Hydrachna, Eylais" (with indication of slide numbers)
1 (1009-1020)
Hydrachna geographica 3
H. leegei 7
H. kisselowi n.sp. (cncelled, "piersigi") 3
H. globosa 2
H. processifera 4
H. ..lugti..[unleserl.] 1
H. sp. Dn 2
2 (987-1008)
H. sp. Dn 2
H. schneideri 1
H. conjecta 3
H. atrata (partly corr. "leegei") 4
H. scutata 1
H. crassipalpis 1
H. thoni 3
H. bimaculata 1
H. geographica 2
H. processifera 2
H. uniscutata 2
3 (965-986)
H. schneideri crassirostris 1
H. valida crassirostris 2
H. sp. 4
H. uniscutata lata 1
H. denudata 1
H. crassa 1
H. atrata 3
H. unidentata 1
H. unidentata pal[..] Thor 1
H. cruenta ("= thoni Piers.") 1
H. breviscuta [!] n. sp. 1
H. comosa palpalis 1
H. thoni 1
H. comosa 1
H. scutata ("= H. thoni") 1
H. schneideri 1
4 (946-963, 995, 1162)
H. uniscutata 2
H. goldfeldi 3
H. sp. 4
H. cruenta 1
H. conjecta 2
H. (aff.) scutata 1
H. schneideri 1
H. valida 1
H. skorikowi 1
H. valida crassirostris 1
H. schneideri crassirostris 2
H. atrata 1
5 (900-920, 933)
Hydryphantes ruber 7
H. ruber prolongata 2
H. octoporus 2
H. sp. 3
H. hellichi (cosinensis) 6
H. dispar 1
H. crassipalpis 1
H. flexuosus 1
6 (1030-1047)
Hydryphantes dispar 5
H. crassipalpis 1
H. bayeri 1
H. dröscheri 2
H. sp. 2
H. kungarensis n. sp. 2
H. planus 1
H. abnormis 1
7 (730-750, without 736, 744, 745, 749)
Eylais setosa 3
E. undulosa 3
E. mülleri 2
E. soari (partim 'anomalia - sp.?') 5
E. sp. 1
E.stagnaliformis 1
E. discreta 1
E. tantilla 1
E. (media Musselius) 6.6.21 '= tullgreni Thor' 1
8 (793-814)
Eylais sp. 22
9 (751-773, all russian specimens, collected about 1920)
Eylais tantilla "Type" 1
E. discreta stagnalis "Type" 1
E. rimosa "Type" 1
E. mutila "typica" 1
E. koenikei "Type" 1
E. tullgreni "Type" 1
E. spinipons "Type" 1
E. mutila "Type" 1
E. extendens "Type" 1
E. thienemanni 1
E. mülleri 2
E. rimosa 1
E. spinipons 1
E. n.sp./sp.n.? 4
10 (774-791, 792 E. undulosa partly destroyed, in loan Gerecke)
E. spinipons 5
E. koenikei 4
E. infundibulifra 8
E. undulosa 1
11 (834-356)
Eylais sp. 2
E. rimosa (1 "= nullipons") 14
E. extendens 7
12 (815-833)
Eylais sp. 18
13 (from here onwards, slides are no more sorted following their number sequence)
Piona coccinea occulta (old label "longicornis") 3
P. coccinea recurva (old label "longicornis") 1
P. longipalpis 1
P. rotunda 5
P. sp. 2
P. controversiosa (partim 'uncata controversiosa') 3
P. rotundoides 1
14 Piona variabilis dispersa 1
P. carnea 2
P. coccinea stjördalensis 1
P. coccinea coccinea 1
P. rotundoides 5
P. sp. 4
P. rotunda 5
P. uncata uncata 1
P. uncata controversiosa 1
P. nodata 1
P. conglobata conjugula 1
P. circularis 2
15 P. nodatoides n. sp. (27.5., 1.6.1921) 2
P. c. stjördalensis 1
P. rotunda 2
P. longicornis 2
P. gyrophora 1
P. nodata inflata 1
P. rotundoides 1
P. nodata lacerata 2
P. sp. 2
P. uncata 2
P. paucipora 1
P. variabilis dispersa 1
P. circularis 2
16 Piona sp. 2
P. disparilis 2
P. variabilis dispersa nov. var. 6
P. nodatoides n. sp. 1
P. coccinea 1
P. variabilis 4
P. longipalpis 1
P. conglobata conjugula 1
P. conglobata conglobata 1
P. rotunda 2
17 P. carnea 1
P. clavicornis 2
P. nodata 2
P. nodata lacerata nov. var. 2
P. nodata inflata nov. var. 1
P. nodatoides 5 (2 "n. sp.")
P. sp. 1
P. conglobata 1
P. variabilis 1
P. latigenitalia R. Marsh. (R. Marsh. leg., dried up) 1 F.
P. circularis 1
P. coccinea 1
P. rotunda 1
P. coccineoides 2
18 P. nodata 1
P. nodatoides n.sp. 1
P. uncata uncata 2
P. rotundoides 1
P. coccinea recurva 3
P. coccinea stjördalensis 1
P. coccinea coccinea 2
P. coccinea occulta 1
P. discrepans scutata nov. var. 1
P. conglobata 2
P. conglobata conjugula 2
P. circularis 1
P. coccineoides 1
P. rotunda 1
19 P. coccinea stjördalensis 3
P. obturbans 1
P. nodata 6
P. coccinea occulta 4
P. nodatoides 1
P. rotundoides 1
P. coccinea coccinea 4
P. coccinea hankensis n. var. 1
P. coccinea recurva 1
20 Eylais tullgreni 6
E. tantilla 1
E. mülleri 1
E. bisinuosa 3
E. sp. 1
E. ussuriensis (setosa uss.) 2
E. hamata 1
E. elliptica 1
E. triarcuata 1
E. tantilliformis 1
E. extendens 1
E. soari 1
E. variabilis 1
21 Hydryphantes affinis (n.sp.) 6
H. clypeatus 4
H. sp. 3
H. ruber prolongatus 2
H. planus 4
H. hellichi 2
22 Eylais sp. 1
E. bisinuosa parviseta 1
E. bisinuosa 6
E. hamata 1
E. shadini 1
E. degenerata 1
E. degenerata asiatica 3
E. tantilliformis n. sp. (8.6.21) 1

10. "Hygrobates Megapus Unionicola Protzia Acercus Neumania Pioninae: Huitfeldtia Hydrochoreutes Pionacercus"
(slippers not in numerical order)
13 (375-391)
Hydrochoreutes ungulatus 5
H. krameri 3
H. sp. 1
Pionacercus leuckarti 4
P. uncinatus 2
Huitfeldtia rectipes 2
2 (449-466)
Neumania limosa 2
N. (triangularis) = deltoides 3
N. callosa 4
N. ambigua "Type" 1
N. angulata n.sp. Type 466 1
2 preps without labels, Nrs 455, 459
7 (501-514, 953)
Acercus scurus 12
A. torris (partim "inflatus", cancelled)
A. sp. (n. sp.) 2
A. bullatus 2
Hygrobates sokolowi Ny. (5.8.1924 Shadin) 1
8 (484-500)
Acercus ornatus 3
A. (Pionopsis) lutescens 10
A. sp. 3
A. ensifer 2
3 (546-563)
Unionicola arcuata (asiatica) 3
U. japonensis 3
U. intermedia 3
U. gracilipalpis 2
U. setipes 3
U. rezvoi (n.sp.) 2
U. crassipes 1
U. sp. 2
4 (394-411)
Megapus nodipalpis robustus n.var. 4
M. nod. fonticolus 3
M. nod. nodipalpis 12
5 (564-583)
Megapus nodipalpis tivdiae 1
M. nod. var., var.II 6
M. nod. constrictus n. var. 6 - among them 576 F. "Cotype"
M. sp. indet. 4
M. gibberipalpis 1
10 (521-539)
Protzia caucasica n. sp. 7
P. indet. (Larva, Schadonoph.) 2
P. eximia 1
P. eximia? "Sok. et Viets det." 1
P. invalvaris, Harz, Kaltetalsbach 12.09.10
Calonyx (cancelled: "Protzia eximia? Viets det."), Harz Riefenbach 27.8.10 1 [= Protzia halberti!]
6 (535-545)
Calonyx sp. 1
C. cabardinicus n. sp. 2
Calonyx flagellum pauciacetabulata 3 - darunter 541 "Typus"
1 (463-483)
Neumania deltoides 3
N. triangularis 1
N. limosa 2
N. sp. 3
N. spinipes 3
N. vernalis 3
N. callosa 1
9 (412-430a)
Megapus ovalis 2
M. sp. 4
M.nodipalpis robustus 1
M. gibberipalpis 4
M. spinipes F. 1
M. spinipes caucasica n. var. 1
M. nodipalpis 2
M. tener 3
11 (430b-448)
Megapus sp. 5
M. arcuatus 3
M. rectipes n. sp. 1
M. gracilis n.sp. 1
M. issajewi n. sp. M Type! 1
M. nodipalpis constrictus 1
M. ovalis 1
N. nodipalpis nodipalpis 1
Neumanisa deltoides 1
12 (584-601)
Hygrobates trigonicus 7
H. fluviatilis 4
H. calliger 3 (2 slides with different labels, Lukin det.)
H. squamifer 1
H. japonicus 1
H. longipalpis Hermann 1
H. bucharicus n.sp. 1
14 (705-722)
Hygrobates longipalpis 10
H. naicus 2
H. nigromaculatus 2
H. trigonicus 2
H. foreli 2
H. sp. "Dictyobates" 1
15 (689-709, 862)
H. fluviatilis 2
H. sp. 1
H. trigonicus 1
H. calliger 3
H. longipalpis 1
H. sokolowi 2
H. sokolowi macrotuberculatus nov var. ("Type") 2
H. nigromaculatus 1
H. neutralis 1
16 (671-686)
H. (Rivobates) n. sp. Type F. 1
H. (Rivobates norvegicus 1
H. Mixobates uncatus 2
H. Mixobates processifer (Archangelsk) 1
H. nigromaculatus 1
H. fluviatilis 1
H. foreli 2
H. sokolowi (Oka 593 m 6.8.24) Ny. 1
Mesobates forcipatus F. 1
17 (651-670)
Hygrobates nigromaculatus 6
H. longipalpis
H. squamifer (partim "?") 11
H. trigonicus 2
18 (637-650)
H. naicus 4
H. fluviatilis 7
H. reticulatus imminutus (...[?] det.)
H. indet. 1
19 (618-636)
H. nigromaculatus 19 (6 "var. octoporus Dad.")
20 (602-617)
H. foreli 16

11. "1"
1 Hydrachna skorikowi skorikowi 1
H. uniscutata sinuata 1
H. Lv. attached to insect tergite 1
2 H. trilobata 2
3 H. comosa Dn 2
4 H. leegei 1
5 H. uniscutata aff. paludosa 2
6 H. aff. processifera 9
7 H. skorikowi skorikowi 4
8 H. globosa 3
H. crassipalpis 2

12. "4"
4 (1233-1298)
Pontarachna anellata n.sp. 1
P. pacifica pilosa n. var. 1
P. indet. 1
Nautarachna crassa F. 2
3 (1198, 1224-47)
egg clusters of: Hydrochoreutes ungulatus, Acercus lutescens, Piona longicornis (?), P. carnea, P. nodatoides, P. variabilis, P. longipalpis, Eylais sp., E. hamata
larvae indet.
Piona Ny.
1 (1179-1190)
Panisoides setipes 1
Thyopsis cancellata (arevalis) 3
Zschokkea oblonga 1
Panisus michaeli 6
Rhypicephales rossica, praeimag. Ixod. 1
2 (1285-1640 - numbers not in order)
Thermacarus thermobius Lv 1
Limnesia fulgida 3
L. undulata 2
L. sp. 1
L. maculata 2
Limnochares aquatica 3
Megapus sp. 2
Halacaridae sp. n. 1
5 (1261.1286)
Georgella sp. 2
Thyas barbigera 11
T. bruzelii 4
T. dirempta 1
Parathyas thoracata 1
Thermohydracarus thermalis Lv 1
6 (1196-1228)
Zschokkea oblonga 1
Sperchon setiger 1
Acercus lutescens Schad. 1
Limnochares aquatica Schad. 1
egg clusters of: Sperchon, Limnesia maculata, L. undulata, Neumania spnipes, Arrenurus neumani, Arrenurus sp., Acercus lutescens
7 (1219-1264)
Thyas pachystoma (1 inepta) 7
T. rivalis (1 colligera) 5
Euthyas truncata Dn 1
Georgella koenikei 2
Hydryphantes (Polyh.) sp. 1
Thyas, eggs with larvae 1
8 (1191-1635 - numbers not in order)
Oxus larvae 1
Oribatei 1
Larvae indet. 3
Diplodontus sp. 1
D. despiciens 2
Neumania spinipes Lv. 1
Piona longicornis Lv. 1
P. variabilis Lv. 1
Thyas sp. Lv. 1
Protzia sp. Lv. 1
Limnesia fulgida Lv. 1
Limnesia maculata Lv. 1
Limnochares aquatica 1
dito Lv. 1

13. "9"
1 Unionicola ypsilophora 6
U. intermedia 9
U. bonzi 2
indet. ? (not decipherable) 1
2 U. ypsilophora 2
U. zervoi [?] 3
3 U. crassipes 12
4 U. ypsilophora 9
U. indet. 2
5 Atractides nodipalpis constrictus 3
Megapus indet. 1
M. aff. nodipalpis robustus 6
M. aff. nodipalpis 4
6 Atractides spinirostris 4
7 A. aff. processifer F. 1
8 Mesobates forcipatus F. 1
9 Megapus ovalis 13
10 Megapus indet. 5
M. subasper 1
11 Torrenticola anomala M. 9
Torrenticola (=Atractides) [in reality Mideopsis orbicularis] 1
Atractides s. str. sp. 1
Atractides (Megapus) F. 1
Atractides s.str. sp. "nymphe" [F.!] 1
12 Atractides sp. [T. barsica-like] 4
Torrenticola amplexa [o.k.] 1

14. "10. Oxus Frontipoda Gnaphiscus"
1 Oxus ovalis 3
Gnaphiscus indet. 1
2 Oxus longisetus 1
3 Frontipoda musculus Dn 1
4 Gnaphiscus ekmani 4
15. "7. Sperchon Sperchonopsis Dartia"
1 Neumania (aff.) spinipes 7
N. deltoides 5
N. limosa 4
N. sp. 3
2 Sperchon glandulosus cubanicus 6
S. plumifer 1
S. clupeifer 1
S. indet. 3
Hygrobates indet. 2
Lebertia (Pilo) indet. 1
3 Sperchon glandulosus 17
4 S. clupeifer 5
S. setiger 1
5 S. papillosus 4
6 S. elegans 4
7 S. squamosus 1
8 S. thori 3
S. undulosus 1
S. indet. 6
9 S. undulosus 5
10 Sperchonopsis verrucosa 2
16. "11. Hydrodroma Thyopsis Thyas"
1 Thyopsis cancellata 3
2 Thyas dirempta 1
T. pachystoma 2
3 "Diplodontus (Eupatra?) sp." [= Huitfeldtia!]
4 Thyas barbigera 4
5 -empty-
6 Hydrodroma (Diplodontus) despiciens 8
"Diplodontus (?) nymphe" [= Pioninae!]
7 Hydrodroma (Diplodontus) despiciens 17
8 dito 20
17. "12. Hydryphantes"
1 Hydryphantes affinis 1
2 H. clypeatus 2
3 H. crassipalpis 1
4 H. octoporus 3
5 H. indet. 4
6 H. ruber 14
7 H. planus 14
H. clypeatus 1
8 H. aff. hellichi 1
9 H. flexuosus 4
Lebertia sp. 1

18. "14. Hydrochoreutes Pionopsis Tiphys Huitfeldtia Forelia Aturus Brachypoda"
1 Forelia liliacea 5
F. sp. 2
2 Hydrochoreutes ungulatus 2
H. krameri 1
3 Tiphys scaurus 3
T. sp. 1
4 T. ornatus 11
5 Acercus ensifer 1
6 Acercus ? F. 1
Tiphys sp. n. (?) M. 1
7 Pionopsis lutescens 13
8 Aturus scaber 2
A. intermedius 2
9 Brachypoda versicolor 10
B. sp. Lv 3
10 Huitfeldtia rectipes 8
19. "15. Ljania Limnesia"
1 Limnesia polonica? 1
2 -empty-
3 Ljania aff. bipapillata 1
4 -empty-
5 Limnesia sp. 10
L. undulata 3
L. aff. connata 1
6 L. maculata 15
L. maculata angustata 3
7 L. maculata 7
8 L. koenikei 4
9 L. undulata 17
10 dito 9
11 dito 16
12 L. fulgida 3
20. "Mongolia"
1 Parasoldanellonyx sop. 4
P. baicalensis 8
2 P. sp. 5
P. baicalensis 9
3 Eylais aff. [not decipherable] 1
E. infundibulifera mongoliensis ssp. n. 2
E. sp. 1
Hydrachna conjecta 1
Hydryphantes dröscheri 2
Hydrodroma despiciens 1
Sperchon sp. 2
S. glandulosus 4
S. tridentatus 2
4 Lebertia jenissajensis 2
L. lepnevae 15
L. beleensis [?] 1
5 L. ussuriensis 3
L. densa 7
L. longissima 8
6 L. ignatovi 8
L. acuta 1
L. sp. 9
7 Oxus angustipositus 1
O. ekmani (also Frontipoda, Gnaphiscus) 3
O. affinis 1
Limnesia fulgida 1
L. undulata
8 Hygrobates octoporus 7
H. foreli 8
9 H. sp. 6
Unionicola crassipes 3
Feltria rubra 2
F. (aff.) minuta 2
Tiphys bullatus 1
Pionopsis lutescens 1
Piona uncata 1
P. rotunda 2
Arrenurus sp. 1
10 -empty-
21. Dsungaria [?]
1-3 -empty-
4 Piona coccinea 1
Arrenurus sp. 3 (1/2/0)
Hydracarina Lv. 1
Halacarae 1

22. "Central Asia Kuzmangalieva leg."
1 Lebertia sp. 3
Sperchon glandulosus ("var.") 7
Feltriidae, Feltria 2
Atractides nodipalpis var.? 1
Feltria minuta 1
Thyas aff. rivalis 1
Sperchon sp. 3
Feltria rubra 2
2 Sperchon sp. 6
Atractides nodipalpis 1
Sperchon squamosus 3
Aturus sp. 2
Feltria minuta 1
F. sp. 1
Lebertia sp. 1
"Partnunia sp. " [Protziella!] 2 (1/1/0)
Atractides sp. 1
- all remaining slippers empty -
23. "Different" (cancelled: Oribatida)
only one slide: Unionicola intermedia 1
24. Kirgisia, Issuk-Kul E shore, River Mun
1, 2 -empty-
3 Sperchon glandulosus cubanicus 7
S. sp. (adults + larvae) 3
4 Hygrobates aff. foreli 2
Megapus nodipalpis fonticolus 9
5 Lebertia (aff.) lineata 6
L. rivulorum 3
L. (aff.) tuberosa 4
L. glabra 3
L. sp. 1
6 Kongsbergia (aff.) materna 6
Aturus duplex 1
larvae indet. 4
Feltria minuta 7
7 Sperchon glandulosus (cubanicus) 9
larvae indet. 2
Sperchon sp. 4
8 Sperchon undulosus /aff. clupeifer 4
S. glandulosus 2
S. sp. 2
Atractides nodipalpis fonticolus 2
A. sp. 1
Feltria minuta 1

25. "Type collection" (species name is followed by slide number[s] and eventual additional indications; in general, if several slide numbers are given, they probably represent syntypes. * = not designated as type by Sokolow)
1 Lebertia exuta okaensis 347
L. tuberosa Kaukasus 193*
L. alonensis 123
L. balcarica 63
L. sutschanika 62 W.
L. acuta 56
L. tschernowskii 49
L. parallelepipeda 45
L. densa aberrans 43
L. asiatica teletzkensis 233*
2 Hydryphantes longicornis 1/638 W.
H. affinis "182" [locality number?] 1847
H. sp. 224, 237 [locality numbers?] *
Megapus sp. 209, 207, 276, 237 [locality numbers?] *
Hygrobates uncatus 674
Lebertia gibberipalpis 1859
L. hissarica 1868, 1858, 1848, 1841
L. zernovi 1844, 1857
3 Unionicola setipes 549
Megapus nodipalpis constrictus 571
Hygrobates bituberosus 593
H. diversiporus 672 (1 loaned to Gerecke)
Mesobates longipes 676
Hygrobates uncatus 683
H. intricatus 687
H. kirgizicus 694, 695
Panisus thori devdorakensis (loaned to Gerecke)
Hydryphantes fonticolus (loaned to Gerecke)
Eylais tantilliformis 757
E. rimsky-korssakowi 760, 761
E. setosa goldfeldi 767
E. setosa ussuriensis 772
Hydrarachna kisselewi 1027
Piona coccinea hankensis 925
4 Lebertia stackelbergi 310
L. beleensis 314
L. caucasia 319 F., 331 Dn
L. lepnevae 343
L. pachypalpis 351 ("Type IV")
Megapus nodipalpis robustus 397 F., 392 M.
M. sansoni 422 M.
M. gracilis 430 ("Type N. IV") F.
M. gassowskii 435 F.
M. odarkensis 439 F.
M. manensis 441 F.
M. manensis rotundus 445 F.
Neumania angulata 467
N. setipalpis 471
N. distans 471 [same slide!]
Calonyx aff. cabardinicus Schadonophan 540 *
5 Lebertia asiatica ralitzkensis 78
L. affinis 65
L. acuta 57, 66
L. shadini 54
L. incurvata F. 47
L. altaica M. 47 [same slide]
L. tschernowskii 46
L. parallelepipeda 40
L. dubiaeformis 38
L. altaica 41
L. jenissejensis 339
L. ignatowi 82
L. ussuriensis 87
L. pireschnikowi 226
L. caucasia cubanica 137
L. altaica (=densa?) 215
L. longissima F. 249, M. 252
L. stackelbergi saxicola 272
L. stackelbergi 305
6 Oxus magnus 1617
Copidognathus derjugini 1639 F., 1641 M.F., 1649 Dn
C. gurjanowae 1687 Lv., 1689 N I
C. lamelliger 1688 NII
Sperchon tarnogradskii 1574
S. minutiporus 1513
S. tridentatus 1526
Rhombognathus seahami rhomboidea 1695
R. denticulatus 1696
R. terminalis 1701
Halacarus mollis 1710
H. pulcher 1718
H. crassidens 1732 NyII, 1738 M.
Agaue alberti caraensis 1741

26. "Lake Baikal" - partly continuation of the type collection!
1 [not decipherable] Lobohalacarus pasgatum [?]
Hydrachna processifera paucipora 1021
Atractides turcestanicus 1863 *
Piona gyrophora (cancelled: azientatis nov sp.) 1127
Pseudosoldanellonyx lohmanelloides Holotype, 5 paratypes
Stigothrombium vermiforme 2 specimens without nr. (8.6.65, 26.7.66)
Lobohalacarus bucharensis Type, Ny.I, Lv., no nr.
L. sp. indet. 1 sp. *
Baicalacarus vermiformis n.sp. n.gen. MM. FF. Dn, 5 prep without nr.
2 Hydrarachna atrata 1192
Hydryphantes? 1194 * no collecting site
Unionicola 1195 *
...Atractides (eggs?) 1197 *
"Hypopus" 1199 *
..Hydrachnellae.. 1201 *
Hydrochoreutes krameri -unleserl.- Schadonophan 1202 *
Thyas barbigera Viets aberr. 1203 *
3 Agauopsis robusta 2 *
Agaue sp. 1 Antarktis 1956 *
Agaue sp. 2 Antarktis 1956 2 preps. *
Halacarus crassidens Antarktis 1956 *
Agaue tenuirostris 4 preps
Bathyhalacarus quadricornis 7 prep. * - but below the (new) labels an original text is visible, with the indication "Paratype" in one specimen. This could be the type series with the designation of the holotype cancelled or fewly visible
4 Parasoldanellonyx parviscutatus 6 preps *
Feltria sp. 1 *
F. minuta 1 *
5 Parasoldanellonyx baicalensis 9
P. sp. 1 *
Soldanellonyx chappuisi 1
6 Parasoldanellonyx baicalensis 9
P. sp. 3
7 Dartia emarginata 1611
Agaue alberti orientalis 1743
Agaue alberti caraensis 1742
Agauopsis robusta 1751 NyII, 1761 NyI
Tadjicothyas connexa 1861
Pontarachna anellata 1296
Rhombognathus aculeatus 1691
8 Thermacarus thermobius F., 1283, 1284 (legs); M. 1291, 1287 (legs)
Rutripalpus limicola F. 1289, 1288
Pontarachna pacifica pilosa 1292
Arrhenurus bucharicus 1308
A. wereschtschagini 1328
A. quadrimaculatus 1344
A. daubinensis 1348
A. ussuriensis 1354
Kondia karelica 1193
Albia stationis orientalis 1410 F.
Aturus complexus 1419
Oxus magnus 1447 Dn
Atractides ussuriensis 1554 "Type N.III" F.
9 Hydryphantes? 1 *
27. Kasachstan, Tien schan, Pamir
1 Forelia liliacea 5
Hydryphantes flexuosus 2
Soldanellonyx chappuisi 1
Limnesia undulata 1
Hygrobates aff. kirgisicus 1
Sperchon clupeifer 1
Eylais rimosa 1
Hydrachna skorikowi 1
2 Trombiidae 2
Gamasidae 1
"Torrenticola elliptica" ["Rusetria"-like!] 0/3/2
Lebertia affinis 2
L. sp. 2
Sperchon glandulosus ssp. 1
3 Dartia borneri 3
D. sp. 3
D. longipora 1 Dn
Calonyx cabardinicus F. 1
Panisus sp. 1
Kongsbergia sp. Dn. 1
Axonopsidae 1
4 Soldanellonyx chappuisi 3
Parasoldanellonyx pauciscutatus 1
5 indet. 3
Halacaridae 1
Lobohalacarus weberi 1
Soldanellonyx chappuisi 14
6 Sperchon indet.
Feltria (?) minuta 1
Piona nodata 2
Limnesia connata 1
L. sp. 1
Piona sp. 1
7 Sperchon sp. 10
Lebertia sp. 1
Hygrobates aff. kirgisicus 1
"Torrenticola" [really Kongsbergia!] 6
Megapus F. 1
Piona Dn 1
8 Sperchon sp. 1
Hygrobates longiporus 1
9 Sperchon (teils?) 12
Hygrobates sp. 1
H. longiporus 6
Piona nodata 1

28. Kisil-Kumil
1 Arrenurus globator 1
Porohalacarus 1
Acari 1
2 Halacaridae 1
Lobohalacarus 2
Dartia sp. Dn 1
D. borneri 2
Lebertia sp. 2
Pionidae/Piona Dn 2

29. Collections from russian lakes 1978, 1979, 1982
23 slippers with 18 slides each, not sorted systematically. Species list:
Eylais extendens
Zschokkea oblonga
Thyas sp.
T. aff. rivalis
Hydrodroma despiciens
Lebertia lepnevae
L. longissima
L. dubiaeformis
L. insignis
L. aff. ignatovi
L. porosa
L. ienissiensis
L. indet. (much!)
Oxus ekmani
Gnaphiscus aff. affinis
G. setosus
Frontipoda sp.
F. musculus
Limnesia koenikei
L. undulata
L. maculata
L. polonica
Hygrobates foreli
H. squamifer
H. octoporus
H. fluviatilis
H. nigromaculatus
H. sp.
Feltria minuta
F. rubra
Neumania angulata
Unionicola sp.
Tiphys bullatus
Huitfeldtia rectipes
Pionacercus leuckarti
Forelia liliacea
Piona rotundoides
P. rotunda
P. uncata
P. coccinea
P. coccinoides
P. conglobata
P. sp.
Hydrochoreutes krameri
Brachypoda versicolor
Neobrachypoda ekmani
Acalyptonotus violaceus
Midea orbiculata
Mideopsis crassipes
M. orbicularis
Arrenurus mülleri
A. ussuriensis
A. distinctus
A. sp.
Hydrachnellae Lv indet.
Halacaridae indet.
Porolohmanella violacea
Soldanellonyx parviscutatus
S. monodi
S. chappuisi
Porohalacarus alpinus
Gamasidae

30. praeimaginal stages [?] - label inside: "Kasach. SSR..."
1 eggs, larvae (+/- engorged), protonymphs indet, 14 slides
2 dito, 16 slides
3 Arrenurus sp. Dn 1
Sperchon sp. Lv 1
Arrenurus sp. F. 1
4 Hygrobates sp. 1
Atractides sp. M. 1
Feltria sp. Dn 1
Piona sp. F. 1
Larvae indet. 8
Acari indet. 4
5 Larvae indet. 3
protonymphs indet. 2
6 Hydracarina eggs, larvae indet. 2
terr. mites 8
Arrenurus indet. Lv. 1
Lebertia indet. Lv
Sperchon indet. Lv. Dn 2
Sperchonopsis Lv 3
7 indet Dn 1
Piona Lv. (?) 2
Arrenurus Lv. 2
Sperchon? Lv. 1
Porolohmanella violacea 1
Stechmückenlarve 1
8 Sperchon Lv. attached to Wilhelmia mediterranea (Turkmen.) 2
Trombidiformes 2
9 Landmilben indet. 9
10 dito 1
Arrenurus sp F. 1
Piona sp. F. 1
Feltria sp. F. 1
Hydrochoreutes sp. F.
11 Hygrobates sp. Lv.
Unionicola intermedia F. from Anodonta piscinalis 1
cf. Unionicola Lv. from Unio pictorum 3
12 Arrenurus sp. Lv.
13 Eylais sp. Ad. 18
14 Sperchon Lv. 1
Nymphe indet. 2
Rhizopoda 1
Cheyleitus eruditus 1
Leptus sp. 1

31. "Hydracarina collection Sokolow 7 Halacaridae"
1 Halacaridae indet. 1
Halacarus balticus 1 Type
Halacarus alpinus 2
Caspihalacarus hyrcanus 2
Halacarus basteri 4
2 Halacarus (?) hexacanthus 7
H. crassidens 8
H. basteri (affinis) 2
H. mollis 1 Type
H. sp. 1
Halacarellus longipes 1 Type
H. sp. 1
3 Copidognathus derjugini 9
sp. n. 3
Copidognathus n. sp.? 1
C. latus 1 Type
C. oxianus 2
C. uschakowi 2
C. gutjanowae 1
4 C. gorbunovi F. n.sp.? 1
5 C. derjugini 3
C. oxianus 4
C. gurjanowae (major) 12 (Type)
C. elongatus F. 1 Type
C. gracilipalpis n. sp. var. 1
6 Rhombognathopsis seahami 2
R. contectus (?) 3
R. nudus 1
R. levis 2
7 Agaue chevreuxi 1
Lohmanella falcata 1
Porolohmanella violacea 2 (1 Schlesien, Viets det.)
8 Agauopsis robusta 10
A. brevipalpis (pontica) 3
9 indet. (partim Soldanellonyx?) 5
Soldanellonyx sp. 2
S. monardi 5
S. chappuisi 7
Parasoldanellonyx parviscutatus 1

32. "Hydracarina Kollektion Sokolow 6, varia
1 Forelia ligulifera 2
F. variegator 1
F. liliacea
F. parmata 2
F. parmata scuifera 1 Type
Dartia longipora 1
Atractides [Torr.] sp. 2
Gnaphiscus ekmani 3 (1 "Type")
G. intermedius 3 (2 "Type")
Megapus eggs 1
2 Oxus (?) quadriporus 4
O. ovalis 5
O. angustipositus 4
O anostomium [? not decipherable]
O. longisetus 2
O. nodigerus 3
O. sp. 2
Forelia liliacea 1
3 Sperchon glandulosus cubanicus 3 (1 Type)
S. gland. thienemanni 6
S. glandulosus 4
S. sp. 2
S. squamosus 1
S. clupeifer 1
4 S. glandulosus thienemanni 6
S. plumifer 1
S. glandulosus cubanicus 1
S. glandulosus 1
S. denticulatus 2
S. brevirostris 1
Forelia curvipalpis 1
F. parmata (mutata, scutigera) 3
F. liliacea 2
F. sp. 1
5 Sperchon plumifer 19 (partim Lukin det.)
S. sp. 1
S. resupinus 1
6 Sperchon clupeifer 4
S. squamosus 5
S. setiger 1
S. resupinus 1
S. denticulatus "F." [M.!] 1
Atractides ellipticus (Ussuri) F. 2
A. amplexus 2
A. connexus 1
A. sandalensis 2 (1 only palps, dried)
7 Sperchon minutiporus 1
S. plumifer 4
S. tarnogradskii 1
Pseudosperchon verrucosus (caucasicus) 4
Sperchon glandulosus 1
S. gland. thienemanni 1
S. gland. cubanicus 1
S clupeifer 1
S. squamosus 2
S. papillosus 1
S. turgidus 2
S. setiger 1
S. sp. 3
8 Sperchon sp. 5
S. tridentatus 2
S. (?) setiger 1
S. plumifer 1
S. papillosus 1
S. denticulatus 1
S. squamosus 5
S. clupeifer 2
Pseudosperchon verrucosus 1
9 Sperchon glandulosus 4
Sp. gland. thienemanni 2
S. denticulatus 1
S. undulosus 1
Atractides ussuriensis 2
A. connexus 1
A. ellipticus 2
A. sp. 2
A. amplexus 2
10 Oxus ovalis
Dartia longipora 1

33. "collection Sokolow 5"
1 Arrenurus tricuspidator M.F. 5
A. caudatus W. 3
A. wereschtschagini M. 1
A. bicuspidator M.F. 2
A. crassicaudatus M. 2
A. sinuator M. 1
A. albator M. 1
A. compactus M. 1
A. sp. F. 3
A. neumanioides F. 1
2 A. albator F. 1
A. daubikensis F. 1
A. nassonowi F.
A. forcipatus M. 1
A. innominatus F. 1
A. soochowensis Marsh., USSR det Sokolow, coll. 7.7.27 M. 1
A. sp. M.F. 2
A. crassipetiolatus W. 1
A. maculator F. 1
A. karelicus M. 1
A. sinuator F. 1
A. affinis ? 1
A. mülleri M. 1
A. distinctus Marsh. F. 1 Cotype
3 A. caudatus F. 1
A. globator F. 1
A. sp. M.F.Dn 6
A. cuspidator F. 1
A. claviger F. 1
A. batillifer M.F. 2
A. bisulcicodulus M. 1
A. forpicatus F. 1
A. affinis F. 1
4 Brachypoda versicolor 4
Xystonotus willmanni 1
Neobrachypoda sp. Lv 1
N. ekmani 1
Axonopsis elongata F. 1 Type
5 Arrenurus (?) neumani F. 3
Brachypoda versicolor 1
Aturus sp. 2
A. ussuriensis F. 1 Type
A. natangensis 3
A. scaber 3
A. latus F. 1 Type
6 Mideopsis orbicularis 8
M. crassipes 2
Midea orbiculata 2
Kongsbergia materna 2
Acalyptonotus violaceus 1
Arrenurus distinctus M. 1 Cotype
7 Feltria rubra 6
F. minuta 2
F. sp. Lv 1
8 Acalyptonotus violaceus 1
Limnesia sp. M.F.Dn 5
L. undulata 5
9 L. fulgida 1
L. undulata 2
L. koenikei 2
L. maculata angustata F. 1 Type
L. maculata 2
L. connata 2
L. sp. 7

34. "Lebertia Teutonia"
1 Lebertia (Pilo) indet. 12
L. (Pilo) "latigenital" 3
L. (Pilo) n.sp. 1 "Type" (no name, Nowaja Semlja 29.8.24)
L. porosa 1
L. (Neol.) indet. 1
L. (Hexa) minutipora 1 Type
Hygrobates Lv. 1
2 Lebertia (Pilo) indet. 10
L. indet. 5
L. diabuschinskii (?) 1
L. porosa 3
3 L. rufipes reticulata 1
L. porosa 4
L. sp. 1
L. (Pilo) sp. 2
L. inaequalis 1
L. caucasica 1
L. insignis 1
L. stackelbergi saxicola 1
L. exuta okaensis 1
L. lineata 1
L. lepnevae 2
L. beleensis 1
4 L. stigmatifera 14
L. sp. 3
L. dubia 2
L. marginata 1
L. sefvei 1
5 L. schmidtii 8
L asiatica teletzkensis (?) 2
6 L. fimbriata 5
L. sp. 3
L. acuta 2 (1 Type)
L. rufipes reticulata 1
L. ignatowi 1
L. stackelbergi saxicola
L. caucasica cubanica 1
L. lineata 1
L. tuberosa 1
7 L. lineata 1
L. minutipalpis 4
L. gladiator 10
L. rufipes 1
L. rufipes reticulata 2
L. rufipes zachariasi 2
L. rivulorum 2
L. sparsicapillata 1
L. dubia 1
L. ussuriensis 1
8 L. porosa 16
L. caucasica 1
9 L. riabuschinskii 1
L. stackelbergi 3
L. stack. saxicola 15
L. caucasica 2
L. insignis 1
10 L. asiatica teletzkanini 1
L. lineata 5
L. longissima 1
L. porosa 1
L. sp. 2
11 L. riabuschinskii 13 (2 "Type", but not from coll. Thor)
L. porosa 1
L. stackelbergi 1
L. stack. saxicola 2
L. (Pilo) n.sp. aff. stackelb. (no name) 1
12 L. korbuensis 1 Type
L. inaequalis 2
L. riabuschinskii 3
L. lepnevae 3
L. exuta okaensis 3
L. saxonica 2
L. porosa 1
L. griebi ("nov. nom. pro aberrans") 2
13 L. lineata 19
14 L. ignatowi 1
L. dubiaeformis 5
L. sp. 2
L. densa 6
L. asiatica teletzkensis 1
L. lepnevae 1 <